WORKLIST ENTRIES (1):

WNT4PROTEIN View alignment      Wnt-4 protein signature
 Type of fingerprint: COMPOUND with 4  elements
Links:
   PRINTS; PR01349 WNTPROTEIN
   PRINTS; PR01841 WNT1PROTEIN; PR01842 WNT2PROTEIN; PR01843 WNT3PROTEIN
   PRINTS; PR01845 WNT6PROTEIN; PR01891 WNT7PROTEIN; PR01892 WNT8PROTEIN
   PRINTS; PR01893 WNT10PROTEIN; PR01894 WNT14PROTEIN; PR01895 WNT16PROTEIN
   MIM; 603490

 Creation date 11-MAR-2003

   1. WODARZ, A. AND NUSSE, R.
   Mechanisms of Wnt signal transduction.
   ANNU.REV.CELL DEV.BIOL. 14 59-88 (1998).

   2. BEJSOVEC, A.
   Signal transduction: Wnt signalling shows its versatility.
   CURR.BIOL. 9 R684-R687 (1999).

   3. DE FERRARI, G.V. AND INESTROSA, N.C.
   Wnt signaling function in Alzheimer's disease.
   BRAIN RES.BRAIN RES.REV. 33 1-12 (2000).
  
   4. SEMENOV, M.V. AND SNYDER, M.
   Human dishevelled genes constitute a DHR-containing multigene family.
   GENOMICS 42 302-310 (1997).

   5. PEIFER, M. AND POLAKIS, P.
   Wnt signalling in oncogenesis and embryogenesis - a look outside the
   nucleus.
   SCIENCE 287 1606-1609 (2000).

   6. MOON, R.T.
   An introduction to non-canonical Wnt and Frizzled signaling.
   SEMIN.CELL DEV.BIOL. 13 215 (2002).

   7. GAVIN, B.J., MCMAHON, J.A. AND MCMAHON, A.P.
   Expression of multiple novel Wnt-1/int-1-related genes during fetal and
   adult mouse development.
   GENES DEV. 4 2319-2332 (1990).

   8. MCMAHON, J.A., MCMAHON, A.P. AND WEINBERG, R.A.
   Essential function of Wnt-4 in mammary gland development downstream of
   progesterone signaling.
   GENES DEV. 14 650-654 (2000).

   9. STARK, K., VAINIO, S., VASSILEVA, G. AND MCMAHON, A.P.
   Epithelial transformation of metanephric mesenchyme in the developing
   kidney regulated by Wnt-4.
   NATURE 372 679-683 (1994).

   Wnt proteins constitute a large family of secreted molecules that are
   involved in intercellular signalling during development. The name derives
   from the first 2 members of the family to be discovered: int-1 (mouse) and
   wingless (Drosophila) [1]. It is now recognised that Wnt signalling controls
   many cell fate decisions in a variety of different organisms, including
   mammals [2]. Wnt signalling has been implicated in tumorigenesis, early
   mesodermal patterning of the embryo, morphogenesis of the brain and kidneys,
   regulation of mammary gland proliferation and Alzheimer's disease [3,4].
  
   Wnt-mediated signalling is believed to proceed initially through binding to
   cell surface receptors of the frizzled family; the signal is subsequently
   transduced through several cytoplasmic components to B-catenin, which enters
   the nucleus and activates the transcription of several genes important in
   development [5]. More recently, however, several non-canonical Wnt
   signalling pathways have been elucidated that act independently of
   B-catenin [6]. Members of the Wnt gene family are defined by their sequence
   similarity to mouse Wnt-1 and Wingless in Drosophila. They encode proteins
   of ~350-400 residues in length, with orthologues identified in several,
   mostly vertebrate, species. Very little is known about the structure of 
   Wnts as they are notoriously insoluble; in terms of primary structure, the
   family is characterised by a signal sequence and an almost invariant pattern
   of 23-24 conserved cysteines [1]. Fifteen major Wnt gene families have been 
   identified in vertebrates, with multiple subtypes within some classes.
  
   Wnt-4 cDNA was isolated from mouse using a PCR-based strategy, where it was
   found to be expressed in adult tissues, particularly in brain and lung [7].
   Wnt-4 is believed to act downstream of progesterone signalling to play
   an important role in mammary gland development [8]. Furthermore, mutations
   in the Wnt-4 gene have been linked to kidney defects [9].
  
   WNT4PROTEIN is a 4-element fingerprint that provides a signature for Wnt-4
   proteins. The fingerprint was derived from an initial alignment of 7
   sequences: the motifs were drawn from conserved regions spanning the N-
   terminal half of the alignment, focusing on those sections that characterise
   Wnt-4 proteins but distinguish them from related Wnt subtypes - motifs 1 and
   2 lie between the putative signal peptide and the first conserved cysteine; 
   and motifs 3 and 4 reside in regions between other well-conserved N-terminal
   cysteines. Two iterations on SPTR40_20f were required to reach convergence,
   at which point a true set comprising 9 sequences was identified.

  SUMMARY INFORMATION
      9 codes involving  4 elements
      0 codes involving  3 elements
      0 codes involving  2 elements

   COMPOSITE FINGERPRINT INDEX
  
    4|   9    9    9    9  
    3|   0    0    0    0  
    2|   0    0    0    0  
   --+---------------------
     |   1    2    3    4  

True positives..
 Q96T81         WNT4_MOUSE     WNT4_HUMAN     WNT4_CHICK     
 WNT4_RAT       WNT4_XENLA     WNT4_BRARE     WNT4_DROME     
 Q9VM29         


  PROTEIN TITLES
   Q96T81           SIGNALING PROTEIN WNT-4 - Homo sapiens (Human).
   WNT4_MOUSE       WNT-4 protein precursor - Mus musculus (Mouse).
   WNT4_HUMAN       WNT-4 protein precursor - Homo sapiens (Human).
   WNT4_CHICK       WNT-4 protein precursor - Gallus gallus (Chicken).
   WNT4_RAT         WNT-4 protein precursor - Rattus norvegicus (Rat).
   WNT4_XENLA       WNT-4 protein precursor (XWNT-4) - Xenopus laevis (African c
   WNT4_BRARE       WNT-4 protein precursor - Brachydanio rerio (Zebrafish) (Zeb
   WNT4_DROME       WNT-4 protein precursor (DWNT-4) - Drosophila melanogaster (
   Q9VM29           WNT4 PROTEIN - Drosophila melanogaster (Fruit fly).

SCAN HISTORY SPTR40_20f 2 300 NSINGLE INITIAL MOTIF SETS WNT4PROTEIN1 Length of motif = 11 Motif number = 1 Wnt-4 protein motif I - 1 PCODE ST INT YLAKLSSVGSI WNT4_XENLA 27 27 YLAKLSSVGSI WNT4_RAT 27 27 YLAKLSSVGSI WNT4_HUMAN 27 27 YLAKLSSVGSI WNT4_MOUSE 27 27 YGGTMLNGGGV WNT4_DROME 32 32 YGGTMLNGGGV Q9VM29 32 32 YLAKLSSVGSI WNT4_CHICK 27 27 WNT4PROTEIN2 Length of motif = 13 Motif number = 2 Wnt-4 protein motif II - 1 PCODE ST INT SEEETCEKLKGPI WNT4_XENLA 38 0 SEEETCEKLKGLI WNT4_RAT 38 0 SEEETCEKLKGLI WNT4_HUMAN 38 0 SEEETCEKLKGLI WNT4_MOUSE 38 0 SNTNNMDMQQGLY WNT4_DROME 54 11 SNTNNMDMQQGLY Q9VM29 54 11 SEEETCEKLKGLI WNT4_CHICK 38 0 WNT4PROTEIN3 Length of motif = 10 Motif number = 3 Wnt-4 protein motif III - 1 PCODE ST INT RRGAQLAIEE WNT4_XENLA 68 17 RHGAQLAIEE WNT4_RAT 68 17 RRGAQLAIEE WNT4_HUMAN 68 17 RRGAQLAIEE WNT4_MOUSE 68 17 RRLATTHCEE WNT4_DROME 117 50 RRLATTHCEE Q9VM29 117 50 RRGAQLAIEE WNT4_CHICK 68 17 WNT4PROTEIN4 Length of motif = 12 Motif number = 4 Wnt-4 protein motif IV - 1 PCODE ST INT TLDTLPVFGKVV WNT4_XENLA 91 13 TLDSLPVFGKVV WNT4_RAT 91 13 TLDSLPVFGKVV WNT4_HUMAN 91 13 TLDSLPVFGKVV WNT4_MOUSE 91 13 TRGKRNIFKKLY WNT4_DROME 139 12 TRGKRNIFKKLY Q9VM29 139 12 TLDTLPVFGKVV WNT4_CHICK 91 13 FINAL MOTIF SETS WNT4PROTEIN1 Length of motif = 11 Motif number = 1 Wnt-4 protein motif I - 2 PCODE ST INT YLAKLSSVGSI Q96T81 27 27 YLAKLSSVGSI WNT4_MOUSE 27 27 YLAKLSSVGSI WNT4_HUMAN 27 27 YLAKLSSVGSI WNT4_CHICK 27 27 YLAKLSSVGSI WNT4_RAT 27 27 YLAKLSSVGSI WNT4_XENLA 27 27 YLAKLSSVGSI WNT4_BRARE 27 27 YGGTMLNGGGV Q9VM29 32 32 YGGTMLNGGGV WNT4_DROME 32 32 WNT4PROTEIN2 Length of motif = 13 Motif number = 2 Wnt-4 protein motif II - 2 PCODE ST INT SEEETCEKLKGLI Q96T81 38 0 SEEETCEKLKGLI WNT4_MOUSE 38 0 SEEETCEKLKGLI WNT4_HUMAN 38 0 SEEETCEKLKGLI WNT4_CHICK 38 0 SEEETCEKLKGLI WNT4_RAT 38 0 SEEETCEKLKGPI WNT4_XENLA 38 0 SDEETCEKLRGLI WNT4_BRARE 38 0 SNTNNMDMQQGLY Q9VM29 54 11 SNTNNMDMQQGLY WNT4_DROME 54 11 WNT4PROTEIN3 Length of motif = 10 Motif number = 3 Wnt-4 protein motif III - 2 PCODE ST INT RRGAQLAIEE Q96T81 68 17 RRGAQLAIEE WNT4_MOUSE 68 17 RRGAQLAIEE WNT4_HUMAN 68 17 RRGAQLAIEE WNT4_CHICK 68 17 RHGAQLAIEE WNT4_RAT 68 17 RRGAQLAIEE WNT4_XENLA 68 17 RRGAQLAIDE WNT4_BRARE 68 17 RRLATTHCEE Q9VM29 117 50 RRLATTHCEE WNT4_DROME 117 50 WNT4PROTEIN4 Length of motif = 12 Motif number = 4 Wnt-4 protein motif IV - 2 PCODE ST INT TLDSLPVFGKVV Q96T81 91 13 TLDSLPVFGKVV WNT4_MOUSE 91 13 TLDSLPVFGKVV WNT4_HUMAN 91 13 TLDTLPVFGKVV WNT4_CHICK 91 13 TLDSLPVFGKVV WNT4_RAT 91 13 TLDTLPVFGKVV WNT4_XENLA 91 13 TLESVPVFGKVV WNT4_BRARE 91 13 TRGKRNIFKKLY Q9VM29 139 12 TRGKRNIFKKLY WNT4_DROME 139 12

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