WORKLIST ENTRIES (1):

MUSCRINICM4R View alignment View Structure    Muscarinic M4 receptor signature
 Type of fingerprint: COMPOUND with 5  elements
Links:
   PRINTS; PR00237 GPCRRHODOPSN; PR00247 GPCRCAMP; PR00248 GPCRMGR
   PRINTS; PR00249 GPCRSECRETIN; PR00250 GPCRSTE2; PR00899 GPCRSTE3
   PRINTS; PR00251 BACTRLOPSIN
   PRINTS; PR00243 MUSCARINICR; PR00538 MUSCRINICM1R; PR00539 MUSCRINICM2R
   PRINTS; PR00540 MUSCRINICM3R; PR00542 MUSCRINICM5R
   INTERPRO; IPR001432

 Creation date 01-JUN-1996; UPDATE 07-JUN-1999

   1. ATTWOOD, T.K. AND FINDLAY, J.B.C. 
   Fingerprinting G protein-coupled receptors.
   PROTEIN ENG. 7(2) 195-203 (1994).

   2. ATTWOOD, T.K. AND FINDLAY, J.B.C. 
   G protein-coupled receptor fingerprints.
   7TM, VOLUME 2, EDS. G.VRIEND AND B.BYWATER (1993).

   3. BIRNBAUMER, L.
   G proteins in signal transduction.
   ANNU.REV.PHARMACOL.TOXICOL. 30 675-705 (1990).

   4. CASEY, P.J. AND GILMAN, A.G.
   G protein involvement in receptor-effector coupling.
   J.BIOL.CHEM. 263(6) 2577-2580 (1988).

   5. ATTWOOD, T.K. AND FINDLAY, J.B.C. 
   Design of a discriminating fingerprint for G protein-coupled receptors.
   PROTEIN ENG. 6(2) 167-176 (1993).

   6. KERLAVAGE, A.R., FRASER, C.M., CHUNG, F-Z. AND VENTER, J.C.
   Molecular structure and evolution of adrenergic and cholinergic receptors.
   PROTEINS 1 287-301 (1986).

   7. WATSON, S. AND ARKINSTALL, S.
   Acetylcholine.
   IN THE G PROTEIN-LINKED RECEPTOR FACTSBOOK, ACADEMIC PRESS, 1994, PP.7-18.

   G protein-coupled receptors (GPCRs) constitute a vast protein family that 
   encompasses a wide range of functions (including various autocrine, para-
   crine and endocrine processes). They show considerable diversity at the 
   sequence level, on the basis of which they can be separated into distinct 
   groups. We use the term clan to describe the GPCRs, as they embrace a group
   of families for which there are indications of evolutionary relationship, 
   but between which there is no statistically significant similarity in 
   sequence [1,2]. The currently known clan members include the rhodopsin-like 
   GPCRs, the secretin-like GPCRs, the cAMP receptors, the fungal mating
   pheromone receptors, and the metabotropic glutamate receptor family.
   
   The rhodopsin-like GPCRs themselves represent a widespread protein family 
   that includes hormone, neurotransmitter and light receptors, all of
   which transduce extracellular signals through interaction with guanine
   nucleotide-binding (G) proteins. Although their activating ligands vary 
   widely in structure and character, the amino acid sequences of the 
   receptors are very similar and are believed to adopt a common structural 
   framework comprising 7 transmembrane (TM) helices [3-5]. 
  
   The muscarinic acetylcholine receptors, present in the central nervous
   system, spinal cord motoneurons and autonomic preganglia, modulate a
   variety of physiological functions, including airway, eye and intestinal
   smooth muscle contractions; heart rate; and glandular secretions. The
   receptors mediate adenylate cyclase attenuation, calcium and potassium
   channel activation, and phosphatidyl inositol turnover [6]. This diversity
   may result from the occurrence of multiple receptor subtypes (of which 5
   are currently known, designated M1 to M5), which have been classified
   based on observed differences in ligand binding to receptors in membranes
   from several tissues.
  
   The M4 receptor is found in high levels in neuronal cells of the CNS [7];
   its distribution largely overlaps with that of M1 and M3 subtypes. It is
   also found in cell lines. No selective agonist has been described [7].
  
   MUSCRINICM4R is a 5-element fingerprint that provides a signature for the
   muscarinic M4 receptors. The fingerprint was derived from an initial
   alignment of 4 sequences: the motifs were drawn from conserved sections
   within the third cytoplasmic loop. Two iterations on OWL28.0 were required
   to reach convergence, at which point a true set comprising 5 sequences was
   identified. Two partial matches were also found (S74916 and ACM4_BOVIN), 
   both of which are M4 receptor fragments.
  
   An update on SPTR37_9f identified a true set of 5 sequences.

  SUMMARY INFORMATION
      5 codes involving  5 elements
      0 codes involving  4 elements
      0 codes involving  3 elements
      0 codes involving  2 elements

   COMPOSITE FINGERPRINT INDEX
  
    5|   5    5    5    5    5  
    4|   0    0    0    0    0  
    3|   0    0    0    0    0  
    2|   0    0    0    0    0  
   --+--------------------------
     |   1    2    3    4    5  

True positives..
 ACM4_RAT       ACM4_MOUSE     ACM4_HUMAN     ACM4_CHICK     
 ACM4_XENLA     


  PROTEIN TITLES
   ACM4_RAT         MUSCARINIC ACETYLCHOLINE RECEPTOR M4 - RATTUS NORVEGICUS (RA
   ACM4_MOUSE       MUSCARINIC ACETYLCHOLINE RECEPTOR M4 - MUS MUSCULUS (MOUSE).
   ACM4_HUMAN       MUSCARINIC ACETYLCHOLINE RECEPTOR M4 - HOMO SAPIENS (HUMAN).
   ACM4_CHICK       MUSCARINIC ACETYLCHOLINE RECEPTOR M4 - GALLUS GALLUS (CHICKE
   ACM4_XENLA       MUSCARINIC ACETYLCHOLINE RECEPTOR M4 - XENOPUS LAEVIS (AFRIC

SCAN HISTORY OWL28_0 2 100 NSINGLE SPTR37_9f 2 6 NSINGLE INITIAL MOTIF SETS MUSCRINICM4R1 Length of motif = 11 Motif number = 1 Muscarinic M4 receptor motif I - 1 PCODE ST INT VHKHRPEGPKE ACM4_HUMAN 226 226 VHKHRPEGPKE ACM4_MOUSE 225 225 VHKHRPEGPKE ACM4_RAT 225 225 VRRHKPESRKE ACM4_CHICK 236 236 MUSCRINICM4R2 Length of motif = 16 Motif number = 2 Muscarinic M4 receptor motif II - 1 PCODE ST INT KKAKTLAFLKSPLMKQ ACM4_HUMAN 237 0 KKAKTLAFLKSPLMKP ACM4_MOUSE 236 0 KKAKTLAFLKSPLMKP ACM4_RAT 236 0 RKGKSLSFFKAPPVKQ ACM4_CHICK 247 0 MUSCRINICM4R3 Length of motif = 21 Motif number = 3 Muscarinic M4 receptor motif III - 1 PCODE ST INT KDTSNESSSGSATQNTKERPA ACM4_HUMAN 290 37 KDTSNESSSGSATQNTKERPP ACM4_MOUSE 289 37 KDTSNESSSGSATQNTKERPP ACM4_RAT 289 37 KETSNESSTVSMTQTTKDKPT ACM4_CHICK 300 37 MUSCRINICM4R4 Length of motif = 19 Motif number = 4 Muscarinic M4 receptor motif IV - 1 PCODE ST INT NPASRWSKIQIVTKQTGNE ACM4_HUMAN 334 23 NPASKWSKIQIVTKQTGSE ACM4_MOUSE 334 24 NPASKWSKIQIVTKQTGNE ACM4_RAT 333 23 NPTSKWSKIKIVTKQTGTE ACM4_CHICK 338 17 MUSCRINICM4R5 Length of motif = 11 Motif number = 5 Muscarinic M4 receptor motif V - 1 PCODE ST INT ANVARKFASIA ACM4_HUMAN 371 18 ANVARKFASIA ACM4_MOUSE 371 18 ANVARKFASIA ACM4_RAT 370 18 ANVARKFASIA ACM4_CHICK 382 25 FINAL MOTIF SETS MUSCRINICM4R1 Length of motif = 11 Motif number = 1 Muscarinic M4 receptor motif I - 2 PCODE ST INT VHKHRPEGPKE ACM4_RAT 225 225 VHKHRPEGPKE ACM4_MOUSE 225 225 VHKHRPEGPKE ACM4_HUMAN 226 226 VRRHKPESRKE ACM4_CHICK 236 236 VRRHCPETRQE ACM4_XENLA 227 227 MUSCRINICM4R2 Length of motif = 16 Motif number = 2 Muscarinic M4 receptor motif II - 2 PCODE ST INT KKAKTLAFLKSPLMKP ACM4_RAT 236 0 KKAKTLAFLKSPLMKP ACM4_MOUSE 236 0 KKAKTLAFLKSPLMKQ ACM4_HUMAN 237 0 RKGKSLSFFKAPPVKQ ACM4_CHICK 247 0 EKKKPISSMKSLLIKQ ACM4_XENLA 237 -1 MUSCRINICM4R3 Length of motif = 21 Motif number = 3 Muscarinic M4 receptor motif III - 2 PCODE ST INT KDTSNESSSGSATQNTKERPP ACM4_RAT 289 37 KDTSNESSSGSATQNTKERPP ACM4_MOUSE 289 37 KDTSNESSSGSATQNTKERPA ACM4_HUMAN 290 37 KETSNESSTVSMTQTTKDKPT ACM4_CHICK 300 37 KETSNESSSASLSHNPPEKQP ACM4_XENLA 290 37 MUSCRINICM4R4 Length of motif = 19 Motif number = 4 Muscarinic M4 receptor motif IV - 2 PCODE ST INT NPASKWSKIQIVTKQTGNE ACM4_RAT 333 23 NPASKWSKIQIVTKQTGSE ACM4_MOUSE 334 24 NPASRWSKIQIVTKQTGNE ACM4_HUMAN 334 23 NPTSKWSKIKIVTKQTGTE ACM4_CHICK 338 17 NTASKWSKIKIVTKQTGNE ACM4_XENLA 334 23 MUSCRINICM4R5 Length of motif = 11 Motif number = 5 Muscarinic M4 receptor motif V - 2 PCODE ST INT ANVARKFASIA ACM4_RAT 370 18 ANVARKFASIA ACM4_MOUSE 371 18 ANVARKFASIA ACM4_HUMAN 371 18 ANVARKFASIA ACM4_CHICK 382 25 VNVARKFASIA ACM4_XENLA 376 23

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