WORKLIST ENTRIES (1):

KV8CHANNEL View alignment       Kv8 potassium channel alpha subunit signature
 Type of fingerprint: COMPOUND with 9  elements
Links:
   PRINTS; PR00169 KCHANNEL; PR01491 KVCHANNEL; PR01495 SHABCHANNEL
   PRINTS; PR01496 SHAKERCHANEL; PR01497 SHALCHANNEL; PR01498 SHAWCHANNEL
   PRINTS; PR01492 KV6CHANNEL; PR01494 KV9CHANNEL

 Creation date 14-MAR-2001

   1. MILLER, C.
   An overview of the potassium channel family.
   GENOME BIOL. 1(4) 1-5 (2000).

   2. ASHCROFT, F.M.
   Voltage-gated K+ channels.
   IN ION CHANNELS AND DISEASE, ACADEMIC PRESS, 2000, PP.97-123.

   3. SANSOM, M.S.
   Putting the parts together.
   CURR.BIOL. 9(19) R738-R741 (1999).   

   4. SALINAS, M., DUPRAT, F., HEURTEAUX, C., HUGNOT, J-P. AND LAZDUNSKI, M.
   New modulatory alpha subunits for mammalian shab K+ channels.
   J.BIOL.CHEM. 272(39) 24371-24379 (1997).

   5. SALINAS, M., DE WEILLE, J., GUILLEMARE, E., LAZDUNSKI, M. AND HUGNOT, J-P.
   Modes of regulation of shab K+ channel activity by the Kv8.1 subunit.
   J.BIOL.CHEM. 272(13) 8774-8780 (1997).

   Potassium ion (K+) channels are a structurally diverse group of proteins
   that facilitate the flow of K+ ions across cell membranes. They are
   ubiquitous, being present in virtually all cell types. Activation of K+
   channels tends to hyperpolarise cells, reducing the membrane's electrical
   resistance, dampening nervous activity. In eukaryotic cells, K+ channels
   are involved in neural signalling and generation of the cardiac rhythm, and
   act as effectors in signal transduction pathways involving G protein-
   coupled receptors (GPCRs). In prokaryotic cells, they play a role in the
   maintenance of ionic homeostasis [1].
    
   Structurally, Kv channels belong to the subfamily of K+ channels whose
   subunits contain 6 transmembrane (TM) domains: these are the voltage-
   gated K+ (Kv) channels, the KCNQ channels, the EAG-like K+ channels and
   3 kinds of Ca2+-activated K+ channels (BK, IK and SK) [2]. All K+
   channels share a characteristic sequence feature: a TMxTVGYG motif
   that resides between the 2 C-terminal membrane spanning helices, and
   forms the K+-selective pore domain [1-2].
   
   The Kv family can be divided into 4 subfamilies on the basis of sequence
   similarity and function: Shaker (Kv1), Shab (Kv2), Shaw (Kv3) and Shal 
   (Kv4). All consist of pore-forming alpha subunits that associate with 
   different types of beta subunit. To form a functional K+ channel pore,
   4 alpha subunits and 4 beta subunits are required. The alpha subunits
   have 6 well-conserved TM domains, a pore loop region and variable length
   N- and C-termini. TM domains 5 and 6, and the loop region, form the K+ 
   channel pore through which the K+ ions pass [3]. 
  
   More recently, 4 new electrically-silent alpha subunits have been cloned:
   Kv5 (KCNF), Kv6 (KCNG), Kv8 and Kv9 (KCNS). These subunits do not themselves
   possess any functional activity, but appear to form heteromeric channels
   with Kv2 subunits, and thus modulate Shab channel activity [4]. When highly
   expressed, they inhibit channel activity, but at lower levels show more
   specific modulatory actions. Coexpression of Kv8 subunits with Shab subunits
   produces altered channel-activation kinetics that do not fit the standard
   model. In addition, channel inactivation shows a shift in voltage-dependence
   toward more negative potentials and a slower time course. TM domain 6 is
   thought to play a key role in these modulatory effects [5].
  
   KV8CHANNEL is a 9-element fingerprint that provides a signature for the Kv8
   potassium channel alpha subunits. The fingerprint was derived from an
   initial alignment of 2 sequences: the motifs were drawn from conserved 
   regions spanning virtually the full alignment length - motifs 1-7 reside in
   the N-terminal cytoplasmic domain; motif 8 spans the C-terminus of TM domain
   6, leading into the cytoplasmic C-terminus; and motif 9 resides in the 
   C-terminal domain. Two iterations on SPTR39_15f were required to reach
   convergence, at which point a true set comprising 3 sequences was identified.

  SUMMARY INFORMATION
      3 codes involving  9 elements
      0 codes involving  8 elements
      0 codes involving  7 elements
      0 codes involving  6 elements
      0 codes involving  5 elements
      0 codes involving  4 elements
      0 codes involving  3 elements
      0 codes involving  2 elements

   COMPOSITE FINGERPRINT INDEX
  
    9|   3    3    3    3    3    3    3    3    3  
    8|   0    0    0    0    0    0    0    0    0  
    7|   0    0    0    0    0    0    0    0    0  
    6|   0    0    0    0    0    0    0    0    0  
    5|   0    0    0    0    0    0    0    0    0  
    4|   0    0    0    0    0    0    0    0    0  
    3|   0    0    0    0    0    0    0    0    0  
    2|   0    0    0    0    0    0    0    0    0  
   --+----------------------------------------------
     |   1    2    3    4    5    6    7    8    9  

True positives..
 P97557         Q9UHJ4         Q60565         


  PROTEIN TITLES
   P97557           NEURONAL POTASSIUM CHANNEL - Rattus norvegicus (Rat).
   Q9UHJ4           NEURONAL POTASSIUM CHANNEL ALPHA SUBUNIT (POTASSIUM CHANNEL 
   Q60565           POTASSIUM CHANNEL KV8.1 - Mesocricetus auratus (Golden hamst

SCAN HISTORY SPTR39_15f 2 5 NSINGLE INITIAL MOTIF SETS KV8CHANNEL1 Length of motif = 15 Motif number = 1 Kv8 potassium channel alpha subunit motif I - 1 PCODE ST INT FCSEGEGEPLALGDC Q60565 30 30 FCSEGEGEPLALGDC Q9UHJ4 26 26 KV8CHANNEL2 Length of motif = 16 Motif number = 2 Kv8 potassium channel alpha subunit motif II - 1 PCODE ST INT GGSRFVLSQQALSCFP Q60565 50 5 GGSRFVLSQQALSCFP Q9UHJ4 46 5 KV8CHANNEL3 Length of motif = 15 Motif number = 3 Kv8 potassium channel alpha subunit motif III - 1 PCODE ST INT PSPLELCDDANPVDN Q60565 89 23 PSPLELCDDANPVDN Q9UHJ4 85 23 KV8CHANNEL4 Length of motif = 14 Motif number = 4 Kv8 potassium channel alpha subunit motif IV - 1 PCODE ST INT GRLHVMEQLCALSF Q60565 124 20 GRLHVMEQLCALSF Q9UHJ4 120 20 KV8CHANNEL5 Length of motif = 19 Motif number = 5 Kv8 potassium channel alpha subunit motif V - 1 PCODE ST INT QEIQYWGIDELSIDSCCRD Q60565 139 1 QEIQYWGIDELSIDSCCRD Q9UHJ4 135 1 KV8CHANNEL6 Length of motif = 14 Motif number = 6 Kv8 potassium channel alpha subunit motif VI - 1 PCODE ST INT RYFRRKELSETLDF Q60565 158 0 RYFRRKELSETLDF Q9UHJ4 154 0 KV8CHANNEL7 Length of motif = 19 Motif number = 7 Kv8 potassium channel alpha subunit motif VII - 1 PCODE ST INT ESQHESEQDFSQGPCPTVR Q60565 179 7 ESQHESEQDFSQGPCPTVR Q9UHJ4 175 7 KV8CHANNEL8 Length of motif = 11 Motif number = 8 Kv8 potassium channel alpha subunit motif VIII - 1 PCODE ST INT SACYFTLKLKE Q60565 436 238 SACYFTLKLKE Q9UHJ4 432 238 KV8CHANNEL9 Length of motif = 14 Motif number = 9 Kv8 potassium channel alpha subunit motif IX - 1 PCODE ST INT YARSIMEMLRLKGR Q60565 476 29 YARSIMEMLRLKGR Q9UHJ4 472 29 FINAL MOTIF SETS KV8CHANNEL1 Length of motif = 15 Motif number = 1 Kv8 potassium channel alpha subunit motif I - 2 PCODE ST INT FCSEGEGEPLALGDC P97557 29 29 FCSEGEGEPLALGDC Q9UHJ4 26 26 FCSEGEGEPLALGDC Q60565 30 30 KV8CHANNEL2 Length of motif = 16 Motif number = 2 Kv8 potassium channel alpha subunit motif II - 2 PCODE ST INT GGSRFVLSQQALSCFP P97557 49 5 GGSRFVLSQQALSCFP Q9UHJ4 46 5 GGSRFVLSQQALSCFP Q60565 50 5 KV8CHANNEL3 Length of motif = 15 Motif number = 3 Kv8 potassium channel alpha subunit motif III - 2 PCODE ST INT PSPLELCDDANPVDN P97557 88 23 PSPLELCDDANPVDN Q9UHJ4 85 23 PSPLELCDDANPVDN Q60565 89 23 KV8CHANNEL4 Length of motif = 14 Motif number = 4 Kv8 potassium channel alpha subunit motif IV - 2 PCODE ST INT GRLHVMEQLCALSF P97557 123 20 GRLHVMEQLCALSF Q9UHJ4 120 20 GRLHVMEQLCALSF Q60565 124 20 KV8CHANNEL5 Length of motif = 19 Motif number = 5 Kv8 potassium channel alpha subunit motif V - 2 PCODE ST INT QEIQYWGIDELSIDSCCRD P97557 138 1 QEIQYWGIDELSIDSCCRD Q9UHJ4 135 1 QEIQYWGIDELSIDSCCRD Q60565 139 1 KV8CHANNEL6 Length of motif = 14 Motif number = 6 Kv8 potassium channel alpha subunit motif VI - 2 PCODE ST INT RYFRRKELSETLDF P97557 157 0 RYFRRKELSETLDF Q9UHJ4 154 0 RYFRRKELSETLDF Q60565 158 0 KV8CHANNEL7 Length of motif = 19 Motif number = 7 Kv8 potassium channel alpha subunit motif VII - 2 PCODE ST INT ESQHESEQDFSQGPCPTVR P97557 178 7 ESQHESEQDFSQGPCPTVR Q9UHJ4 175 7 ESQHESEQDFSQGPCPTVR Q60565 179 7 KV8CHANNEL8 Length of motif = 11 Motif number = 8 Kv8 potassium channel alpha subunit motif VIII - 2 PCODE ST INT SACYFTLKLKE P97557 435 238 SACYFTLKLKE Q9UHJ4 432 238 SACYFTLKLKE Q60565 436 238 KV8CHANNEL9 Length of motif = 14 Motif number = 9 Kv8 potassium channel alpha subunit motif IX - 2 PCODE ST INT YARSIMEMLRLKGR P97557 475 29 YARSIMEMLRLKGR Q9UHJ4 472 29 YARSIMEMLRLKGR Q60565 476 29

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