WORKLIST ENTRIES (1):

CALCITONINR View alignment      Calcitonin receptor signature
 Type of fingerprint: COMPOUND with 6  elements
Links:
   PRINTS; PR00237 GPCRRHODOPSN; PR00247 GPCRCAMP; PR00248 GPCRMGR
   PRINTS; PR00249 GPCRSECRETIN; PR00250 GPCRSTE2; PR00899 GPCRSTE3
   PRINTS; PR00251 BACTRLOPSIN
   PRINTS; PR01350 CTRFAMILY; PR01351 CGRPRECEPTOR
   INTERPRO; IPR001688

 Creation date 15-APR-1995; UPDATE 06-JUN-1999

   1. ATTWOOD, T.K. AND FINDLAY, J.B.C.
   Fingerprinting G protein-coupled receptors.   
   PROTEIN ENG. 7(2) 195-203 (1994).

   2. ISHIHARA T., NAKAMURA S., KAZIRO, Y., TAKAHASHI, T., TAKAHASHI, K.
   AND NAGATA, S.  
   Molecular cloning and expression of a cDNA encoding the secretin receptor.
   EMBO J. 10 1635-1641 (1991).
  
   3. LIN, H.Y., HARRIS, T.L., FLANNERY, M.S., ARUFFO, A., KAJI, E.H.,
   GORN, A., KOLAKOWSKI, L.F., LODISH, H.F. AND GOLDRING, S.R.
   Expression cloning of adenylate cyclase-coupled calcitonin receptor.
   SCIENCE 254 1022-1024 (1991). 

   4. JUEPPNER, H., ABOU-SAMRA, A.-B., FREEMAN, M., KONG, X.F.,
   SCHIPANI, E., RICHARDS, J., KOLALOWSKI, L.F., HOCK, J., POTTS, J.T.,
   KRONENBERG, H.M. AND SEGRE, G.E.
   A G protein linked receptor for parathyroid hormone and parathyroid
   hormone-related peptide.
   SCIENCE 254 1024-1026 (1991).

   5. ISHIHARA, T., SHIGEMOTO, R., MORI, K., TAKAHASHI, K. AND NAGATA, S.
   Functional expression and tissue distribution of a novel receptor for
   vasoactive intestinal polypeptide.
   NEURON 8(4) 811-819 (1992).

   6. WATSON, S. AND ARKINSTALL, S.
   Calcitonin.
   IN THE G PROTEIN-LINKED RECEPTOR FACTSBOOK, ACADEMIC PRESS, 1994, PP.74-76.

   7. NJUKI, F., NICHOLL, C.G., HOWARD, A., MAK, J.C., BARNES, P.J.,
   GIRGIS, S.I. AND LEGON, S.A.
   A new calcitonin-receptor-like sequence in rat pulmonary blood vessels.
   CLIN.SCI. 85(4) 385-388 (1993).

   G protein-coupled receptors (GPCRs) constitute a vast protein family that   
   encompasses a wide range of functions (including various autocrine, para-   
   crine and endocrine processes). They show considerable diversity at the   
   sequence level, on the basis of which they can be separated into distinct   
   groups. We use the term clan to describe the GPCRs, as they embrace a group
   of families for which there are indications of evolutionary relationship,   
   but between which there is no statistically significant similarity in   
   sequence [1]. The currently known clan members include the rhodopsin-like   
   GPCRs, the secretin-like GPCRs, the cAMP receptors, the fungal mating   
   pheromone receptors, and the metabotropic glutamate receptor family.       
  
   The secretin-like GPCRs include secretin [2], calcitonin [3], parathyroid   
   hormone/parathyroid hormone-related peptides [4] and vasoactive intestinal
   peptide [5], all of which activate adenylyl cyclase and the phosphatidyl-   
   inositol-calcium pathway. The amino acid sequences of the receptors contain
   high proportions of hydrophobic residues grouped into 7 domains, in a manner
   reminiscent of the rhodopsins and other receptors believed to interact with
   G proteins. However, while a similar 3D framework has been proposed to
   account for this, there is no significant sequence identity between these
   families: the secretin-like receptors thus bear their own unique '7TM'
   signature.      
  
   The major physiological role of calcitonin is to inhibit bone resorption
   thereby leading to a reduction in plasma Ca++ [6]. Further, it enhances 
   excretion of ions in the kidney, prevents absorption of ions in the 
   intestine, and inhibits secretion in endocrine cells (e.g. pancreas and
   pituitary). In the CNS, calcitonin has been reported to be analgesic
   and to suppress feeding and gastric acid secretion. It is used to treat
   Paget's disease of the bone. Calcitonin receptors are found predominantly
   on osteoclasts or on immortal cell lines derived from these cells. It is
   found in lower amounts in the brain (e.g. in hypothalamus and pituitary
   tissues) and in peripheral tissues (e.g. testes, kidney, liver and
   lymphocytes). It has also been described in lung and breast cancer cell
   lines. The predominant signalling pathway is activation of adenylyl cyclase
   through Gs, but calcitonin has also been described to have both stimulatory
   and inhibitory actions on the phosphoinositide pathway.
  
   CALCITONINR is a 6-element fingerprint that provides a signature for the
   calcitonin receptors. The fingerprint was derived from an initial alignment
   of 6 sequences: the motifs were drawn from conserved sections within either
   loop or TM regions, focusing on those areas of the alignment that
   characterise the calcitonin receptors but distinguish them from the rest
   of the secretin-like family - motifs 1-3 were drawn from the N-terminal
   region leading into the first TM domain; motif 4 lies at the C-terminus of
   the second TM domain following into the loop region; motif 5 is N-terminal
   to the seventh TM region; and motif 6 was drawn from the C-terminus. Two
   iterations on OWL25.2 were required to reach convergence, at which point a
   true set comprising 9 sequences was identified. A single partial match was
   also found, RNCLR, a new calcitonin-like receptor from rat pulmonary blood
   vessels [7].
  
   An update on SPTR37_9f identified a true set of 6 sequences.

  SUMMARY INFORMATION
      6 codes involving  6 elements
      0 codes involving  5 elements
      0 codes involving  4 elements
      0 codes involving  3 elements
      0 codes involving  2 elements

   COMPOSITE FINGERPRINT INDEX
  
    6|   6    6    6    6    6    6  
    5|   0    0    0    0    0    0  
    4|   0    0    0    0    0    0  
    3|   0    0    0    0    0    0  
    2|   0    0    0    0    0    0  
   --+-------------------------------
     |   1    2    3    4    5    6  

True positives..
 CALR_RAT       CALR_RABIT     CALR_HUMAN     CALR_MOUSE     
 CALR_CAVPO     CALR_PIG       


  PROTEIN TITLES
   CALR_RAT         CALCITONIN RECEPTOR PRECURSOR (CT-R) (C1A/C1B) - RATTUS NORV
   CALR_RABIT       CALCITONIN RECEPTOR PRECURSOR (CT-R) - ORYCTOLAGUS CUNICULUS
   CALR_HUMAN       CALCITONIN RECEPTOR PRECURSOR (CT-R) - HOMO SAPIENS (HUMAN).
   CALR_MOUSE       CALCITONIN RECEPTOR PRECURSOR (CT-R) - MUS MUSCULUS (MOUSE).
   CALR_CAVPO       CALCITONIN RECEPTOR PRECURSOR (CT-R) - CAVIA PORCELLUS (GUIN
   CALR_PIG         CALCITONIN RECEPTOR PRECURSOR (CT-R) - SUS SCROFA (PIG).

SCAN HISTORY OWL25_2 2 120 NSINGLE SPTR37_9f 2 8 NSINGLE INITIAL MOTIF SETS CALCITONINR1 Length of motif = 19 Motif number = 1 Calcitonin receptor motif I - 1 PCODE ST INT LDQEPFLYLVGRKKLLDAQ CLRA_RAT 34 34 LDQEPFLYLVGRKKLLDAQ CLRB_RAT 34 34 LDQEPFLYLVGRKKLLDAQ RATCR 34 34 LDQEPFLYLVGRKKLLDAQ RATCR1B 34 34 IEPKPFLYVVGRKKMMDAQ CALR_HUMAN 34 34 LEPEPFLYILGKQRMLEAQ CALR_PIG 35 35 CALCITONINR2 Length of motif = 18 Motif number = 2 Calcitonin receptor motif II - 1 PCODE ST INT KCYDRIQQLPPYEGEGPY CLRA_RAT 54 1 KCYDRIQQLPPYEGEGPY CLRB_RAT 54 1 KCYDRIQQLPPYEGEGPY RATCR 54 1 KCYDRIQQLPPYEGEGPY RATCR1B 54 1 KCYDRMQQLPAYQGEGPY CALR_HUMAN 54 1 RCYDRMQKLPPYQGEGLY CALR_PIG 55 1 CALCITONINR3 Length of motif = 15 Motif number = 3 Calcitonin receptor motif III - 1 PCODE ST INT TPDKLHNAYVLYYLA CLRA_RAT 138 66 TPDKLHNAYVLYYLA CLRB_RAT 138 66 TPDKLHNAYVSYYLA RATCR 138 66 TPDKLHNAYVSYYLA RATCR1B 138 66 TPEKLKNAYVLYYLA CALR_HUMAN 138 66 TPDKLQNAYILYYLA CALR_PIG 139 66 CALCITONINR4 Length of motif = 15 Motif number = 4 Calcitonin receptor motif IV - 1 PCODE ST INT LVEVVPNGDLVRRDP CLRA_RAT 202 49 LVEVVPNGDLVRRDP CLRB_RAT 202 49 LVEVVPNGDLVRRDP RATCR 202 49 LVEVVPNGDLVRRDP RATCR1B 202 49 LVEVVPNGELVRRDP CALR_HUMAN 218 65 LVVIVPNGELVKRDP CALR_PIG 203 49 CALCITONINR5 Length of motif = 18 Motif number = 5 Calcitonin receptor motif V - 1 PCODE ST INT SNKVLGKIYDYLMHSLIH CLRA_RAT 364 147 SNKVLGKIYDYLMHSLIH CLRB_RAT 401 184 SNKVLGKIYDYLMHSLIH RATCR 364 147 SNKVLGKIYDYLMHSLIH RATCR1B 401 184 SNKMLGKIYDYVMHSLIH CALR_HUMAN 380 147 STPLLGKIYDYVVHSLIH CALR_PIG 365 147 CALCITONINR6 Length of motif = 23 Motif number = 6 Calcitonin receptor motif VI - 1 PCODE ST INT AVAFAEPGGLPIYICHQEPRNPP CLRA_RAT 433 51 AVAFAEPGGLPIYICHQEPRNPP CLRB_RAT 470 51 AVAFAEPGGRPIYICHQEPRNPP RATCR 432 50 AVAFAEPGGRPIYICHQEPRNPP RATCR1B 469 50 AAAAAEAGDIPIYICHQEPRNEP CALR_HUMAN 445 47 AAALAETVEIPVYICHQEPREEP CALR_PIG 434 51 FINAL MOTIF SETS CALCITONINR1 Length of motif = 19 Motif number = 1 Calcitonin receptor motif I - 2 PCODE ST INT LDQEPFLYLVGRKKLLDAQ CALR_RAT 34 34 VEPEPFLYVIGRKKLMDAQ CALR_RABIT 34 34 IEPKPFLYVVGRKKMMDAQ CALR_HUMAN 34 34 LDQEPFLYLVGRKKLLDAQ CALR_MOUSE 34 34 FEPEPYLYSVGRKKLVDAQ CALR_CAVPO 34 34 LEPEPFLYILGKQRMLEAQ CALR_PIG 35 35 CALCITONINR2 Length of motif = 18 Motif number = 2 Calcitonin receptor motif II - 2 PCODE ST INT KCYDRIQQLPPYEGEGPY CALR_RAT 54 1 KCYDRMEQLPPYQGEGPY CALR_RABIT 54 1 KCYDRMQQLPAYQGEGPY CALR_HUMAN 54 1 KCYDRIHQLPSYEGEGLY CALR_MOUSE 54 1 RCYDRMQQLPPYEGEGPY CALR_CAVPO 54 1 RCYDRMQKLPPYQGEGLY CALR_PIG 55 1 CALCITONINR3 Length of motif = 15 Motif number = 3 Calcitonin receptor motif III - 2 PCODE ST INT TPDKLHNAYVLYYLA CALR_RAT 138 66 TPEKLQNAYVLYYLA CALR_RABIT 138 66 TPEKLKNAYVLYYLA CALR_HUMAN 138 66 TSEKLQNAYVLYYLA CALR_MOUSE 138 66 TPEKLQNAYVLYYLA CALR_CAVPO 138 66 TPDKLQNAYILYYLA CALR_PIG 139 66 CALCITONINR4 Length of motif = 15 Motif number = 4 Calcitonin receptor motif IV - 2 PCODE ST INT LVEVVPNGDLVRRDP CALR_RAT 202 49 LVEVVPNGELVRRDP CALR_RABIT 202 49 LVEVVPNGELVRRDP CALR_HUMAN 218 65 LVEVVPNGDLVRRDP CALR_MOUSE 202 49 LVEVVPNGELVRKDP CALR_CAVPO 202 49 LVVIVPNGELVKRDP CALR_PIG 219 65 CALCITONINR5 Length of motif = 18 Motif number = 5 Calcitonin receptor motif V - 2 PCODE ST INT SNKVLGKIYDYLMHSLIH CALR_RAT 401 184 SNKILGKIYDYLMHSLIH CALR_RABIT 364 147 SNKMLGKIYDYVMHSLIH CALR_HUMAN 380 147 SNKVLGKIYDYLMHSLIH CALR_MOUSE 401 184 SNKVLGKIYDYFMHSLIH CALR_CAVPO 364 147 STPLLGKIYDYVVHSLIH CALR_PIG 381 147 CALCITONINR6 Length of motif = 23 Motif number = 6 Calcitonin receptor motif VI - 2 PCODE ST INT AVAFAEPGGLPIYICHQEPRNPP CALR_RAT 470 51 TAASAEEGGIPVYIYHQEPRNDP CALR_RABIT 429 47 AAAAAEAGDIPIYICHQEPRNEP CALR_HUMAN 445 47 AVAFAEPDGLPIYICHQEPRNPP CALR_MOUSE 469 50 AAAEAGGDNIPVYICHQEPRNDP CALR_CAVPO 433 51 AAALAETVEIPVYICHQEPREEP CALR_PIG 450 51

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